Macroalgae N, Seagrass N, Halophila N, Coral host N

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    Macroalgae N

    Concentration of nitrogen biomass per m2 of macrolagae. Macroalgae (or seaweed) grows above all other benthic plants (corals, seagrasses, benthic microalgae). It is parameterised as a non-calcifying leafy algae, with a C:N:P ratio of 550:30:1, and a formulation for calculating the percentage of the bottom covered as 1-exp(-ΩMA MA). In the model, in the absence of both calcifying macroalgae (particularly Halimeda) and unicellular epiphytes, macroalgae represents the biomass of all seaweeds and epiphytes. Light is accessed in the following order: Macroalgae, Seagrass, Coral.

    Seagrass N

    Concentration of nitrogen biomass per m2 of a seagrass form parameterised to be similar to Zostera. This form captures light after it has passed through macroalgae and before it passes through Halophila. This form is better adapted to high light, low nutrient conditions than Halophila as a result of a deeper root structure and being able to shade it. See macroalgae for elemental ratio and bottom cover. Light is accessed in the following order: Macroalgae, Seagrass, Coral.

    Halophila N

    Concentration of nitrogen biomass per m2 of a seagrass form parameterised to be similar to Halophila. This form captures light after it has passed through the Zostera seagrass form. The Halophila form is batter adapted to low light conditions than Zostera, having a faster growth rate and lower minimum light requirement. See macroalgae for elemental ratio and bottom cover. Light is accessed in the following order: Macroalgae, Seagrass, Coral.

    Coral host N

    Concentration of nitrogen biomass per m2 of coral host tissue in the entire grid cell. Unlike other epibenthic variables, corals area is assumed to exist in communities that are potentially smaller than the grid size. The fraction of the grid cell covered by corals is given by ACH. Thus the biomass in the occupied region is given by CH/ACH. The percent coverage of the coral of the bottom for the whole cell is given by ACH (1 - exp(-ΩCH CH/ACH)). With only one type of coral resolved, CH represents the biomass of all symbiotic corals. Since the model contains no other benthic filter-feeders, CH best represent the sum of the biomass of all symbiotic filter-feeding organisms such as corals, sponges, clams etc. C:N:P is 106:16:1. Light is accessed in the following order: Macroalgae, Seagrass, Coral.